Pangaea

Pangaea, often shortened to just Pangea, was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from earlier continental units approximately 325 million years ago, and began to break apart about 175 million years ago. In contrast to the present Earth and its distribution of continental mass, Pangaea was centred on the Equator and surrounded by the superocean Panthalassa. Pangaea is the most recent supercontinent to have existed and the first to be reconstructed by geologists.

Formation
Pangaea is only the most recent supercontinent identified in the geologic record. The forming of supercontinents and their breaking up appears to have been cyclical through Earth's history. There may have been several others before Pangaea.

Paleomagnetic measurements help geologists determine the latitude and orientation of ancient continental blocks, and newer techniques may help determine longitudes. Paleontology helps determine ancient climates, confirming latitude estimates from paleomagnetic measurements, and the distribution of ancient forms of life provides clues on which continental blocks were close to each other at particular geological moments. However, reconstructions of continents prior to the breakup of Pangea, including the ones in this section, remain partially speculative, and different reconstructions will differ in some details.

The fourth-last supercontinent, called Columbia or Nuna, appears to have assembled in the period 2.0–1.8 billion years ago (Ga). Columbia/Nuna broke up and the next supercontinent, Rodinia, formed from the accretion and assembly of its fragments. Rodinia lasted from about 1.3 Ga until about 750 million years ago, but its exact configuration and geodynamic history are not nearly as well understood as those of the later supercontinents, Pannotia and Pangaea.

According to one reconstruction, when Rodinia broke up, it split into four pieces: Laurentia, Baltica, the supercontinent of Paleogondwana, and the smaller Congo craton. Proto-Laurasia and Proto-Gondwana were separated by the Proto-Tethys Ocean. Next Proto-Laurasia itself split apart to form the continents of Laurentia, Siberia, and Baltica. Baltica moved to the east of Laurentia, and Siberia moved northeast of Laurentia. The splitting also created two new oceans, the Iapetus Ocean and Paleoasian Ocean. Most of the above masses coalesced again to form the relatively short-lived supercontinent of Pannotia. This supercontinent included large amounts of land near the poles and, near the equator, only a relatively small strip connecting the polar masses. Pannotia lasted until 540 Ma, near the beginning of the Cambrian period and then broke up, giving rise to the continents of Laurentia, Baltica, and the southern supercontinent of Gondwana.

In the Cambrian period, the continent of Laurentia, which would later become North America, sat on the equator, with three bordering oceans: the Panthalassic Ocean to the north and west, the Iapetus Ocean to the south, and the Khanty Ocean to the east. In the Earliest Ordovician, around 480 Ma, the microcontinent of Avalonia – a landmass incorporating fragments of what would become eastern Newfoundland, the southern British Isles, and parts of Belgium, northern France, Nova Scotia, New England, South Iberia, and northwest Africa – broke free from Gondwana and began its journey to Laurentia. Baltica, Laurentia, and Avalonia all came together by the end of the Ordovician to form a landmass called Euramerica, closing the Iapetus Ocean. The collision also resulted in the formation of the northern Appalachians. Siberia sat near Euramerica, with the Khanty Ocean between the two continents. While all this was happening, Gondwana drifted slowly towards the South Pole. This was the first step of the formation of Pangaea.

The second step in the formation of Pangaea was the collision of Gondwana with Euramerica. By the middle of the Silurian, 430 Ma, Baltica had already collided with Laurentia, forming Euramerica, an event called the Caledonian orogeny. Avalonia had not yet collided with Laurentia, but as Avalonia inched towards Laurentia, the seaway between them, a remnant of the Iapetus Ocean, was slowly shrinking. Meanwhile, southern Europe broke off from Gondwana and began to move towards Euramerica across the Rheic Ocean. It collided with southern Baltica in the Devonian.

By the late Silurian, Annamia and South China split from Gondwana and started to head northward, shrinking the Proto-Tethys Ocean in their path and opening the new Paleo-Tethys Ocean to their south. In the Devonian Period, Gondwana itself headed towards Euramerica, causing the Rheic Ocean to shrink. In the Early Carboniferous, northwest Africa had touched the southeastern coast of Euramerica, creating the southern portion of the Appalachian Mountains, the Meseta Mountains, and the Mauritanide Mountains, an event called the Variscan orogeny. South America moved northward to southern Euramerica, while the eastern portion of Gondwana (India, Antarctica, and Australia) headed toward the South Pole from the equator. North and South China were on independent continents. The Kazakhstania microcontinent had collided with Siberia. (Siberia had been a separate continent for millions of years since the deformation of the supercontinent Pannotia in the Middle Carboniferous.)

The Variscan orogeny raised the Central Pangean Mountains, which were comparable to the modern Himalayas in scale. With Pangea now stretching from the South Pole across the equator and well into the Northern Hemisphere, an intense monsoon climate was established, except for a perpetually wet zone immediately around the central mountains.

Life
Pangaea existed as a supercontinent for 160 million years, from its assembly around 335 million years ago (Early Carboniferous) to its breakup 175 million years ago (Middle Jurassic). During this interval, important developments in the evolution of life took place. The seas of the Early Carboniferous were dominated by rugose corals, brachiopods, bryozoans, sharks, and the first bony fish. Life on land was dominated by lycopsid forests inhabited by insects and other arthropods and the first tetrapods. By the time Pangaea broke up, in the Middle Jurassic, the seas swarmed with molluscs (particularly ammonites), ichthyosaurs, sharks and rays, and the first ray-finned bony fishes, while life on land was dominated by forests of cycads and conifers in which dinosaurs flourished and in which the first true mammals had appeared.

The evolution of life in this interval of time reflected conditions created by the assembly of Pangaea. The assembly of most of the continental crust into one landmass reduced the extent of sea coasts. Increased erosion from uplifted continental crust increased the importance of floodplain and delta environments relative to shallow marine environments. Continental assembly and uplift also meant an increasingly arid climate over much of the surface of the Earth. This favored the evolution of amniotes and seed plants, whose eggs and seeds were better adapted to dry climates. The early drying trend was most pronounced in western Pangaea, which became an epicenter for the evolution and geographical spread of amniotes.

Coal swamps are typically a feature of perpetually wet regions close to the equator. The assembly of Pangaea disrupted the intertropical convergence zone and created an extreme monsoon climate that reduced the deposition of coal to its lowest level in the last 300 million years. During the Permian, coal deposition was largely restricted to the North and South China microcontinents, which were among the few areas of continental crust that had not joined with Pangaea. The extremes of climate conditions in the interior of Pangaea are reflected in bone growth patterns of pareiasaurs and in the growth patterns in gymnosperm forests. Early Triassic Lystrosaurus fossil from South Africa

The lack of oceanic barriers is thought to have favored cosmopolitanism, in which species show a wide geographical distribution. Cosmopolitanism was also driven by mass extinctions, including the Permian–Triassic extinction event, the most severe in the fossil record, and also the Triassic–Jurassic extinction event. These events resulted in disaster fauna showing little diversity and high cosmopolitanism. These include Lystrosaurus, which opportunistically spread to every corner of Pangaea following the Permian-Triassic extinction event. On the other hand, there is evidence that many Pangaean species were provincial, with a limited geographical range, in spite of the lack of geographical barriers. This may be due to the strong variations in climate by latitude and season produced by the extreme monsoon climate. For example, cold-adapted pteridosperms (early seed plants) of Gondwana were blocked from spreading throughout Pangaea by the warming climate, and northern pteridosperms ended up dominating Gondwana in the Triassic.

The tectonics and geography of Pangaea may have worsened the Permian-Triassic extinction event or other extinctions. For example, the reduced area of continental shelf environments may have left marine species vulnerable to extinction. However, no evidence for a species-area effect has been found in more recent and better characterized portions of the geologic record. Another possibility is that reduced sea-floor spreading associated with the formation of Pangaea, and the resulting cooling and subsidence of oceanic crust, may have reduced the number of islands that could have served as refugia for marine species. Species diversity may have already been reduced prior to mass extinction events due to mingling of species possible when formerly separate continents were merged. However, there is strong evidence that climate barriers continued to separate ecological communities in different parts of Pangaea. The eruptions of the Emeishan Traps may have eliminated South China, one of the few continental areas not merged with Pangaea, as a refugium.

Breakup
The first phase began in the Early-Middle Jurassic (about 175 Ma), when Pangaea began to rift from the Tethys Ocean in the east to the Pacific in the west. The rifting that took place between North America and Africa produced multiple failed rifts. One rift resulted in a new ocean, the North Atlantic Ocean.

The Atlantic Ocean did not open uniformly; rifting began in the north-central Atlantic. The South Atlantic did not open until the Cretaceous when Laurasia started to rotate clockwise and moved northward with North America to the north, and Eurasia to the south. The clockwise motion of Laurasia led much later to the closing of the Tethys Ocean and the widening of the "Sinus Borealis", which later became the Arctic Ocean. Meanwhile, on the other side of Africa and along the adjacent margins of east Africa, Antarctica and Madagascar, new rifts were forming that would lead to the formation of the southwestern Indian Ocean that would open up in the Cretaceous.

The second major phase in the break-up of Pangaea began in the Early Cretaceous (150–140 Ma), when the landmass of Gondwana separated into multiple continents (Africa, South America, India, Antarctica, and Australia). The subduction at Tethyan Trench probably caused Africa, India and Australia to move northward, causing the opening of a "South Indian Ocean". In the Early Cretaceous, Atlantica, today's South America and Africa, finally separated from eastern Gondwana (Antarctica, India and Australia). Then in the Middle Cretaceous, Gondwana fragmented to open up the South Atlantic Ocean as South America started to move westward away from Africa. The South Atlantic did not develop uniformly; rather, it rifted from south to north.

Also, at the same time, Madagascar and India began to separate from Antarctica and moved northward, opening up the Indian Ocean. Madagascar and India separated from each other 100–90 Ma in the Late Cretaceous. India continued to move northward toward Eurasia at 15 centimeters (6 in) a year (a plate tectonic record), closing the eastern Tethys Ocean, while Madagascar stopped and became locked to the African Plate. New Zealand, New Caledonia and the rest of Zealandia began to separate from Australia, moving eastward toward the Pacific and opening the Coral Sea and Tasman Sea.